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#620. Mutants were ~70 % of CI machine time. 1. mutation-gate.yml: a green full run (aggregator verified all six shards) leaves a cache marker keyed by material tree + workflow SHA. A scheduled night with the same tree and workflow, marker not older than 7 days, skips the shards and writes "reused from run N" to the run summary. Red runs leave no marker, so the next night runs again and still files the issue (#472). Manual dispatch always runs the full registry. Decision is a pure function in scripts/mutation-nightly-reuse.mjs. 2. changed_mutants: the shard plan (already computed before setup, #518) now names the environment of its guards (plan-browser=/plan-python=, from scripts/mutation-environment.mjs). Python + backend deps only for shards with pytest guards, Chromium only for shards whose guards reach Playwright; pip wheels cached. Every shard still runs and reports, so the six mutant jobs of the review proof are unchanged. Item 3 of the issue (smoke guards -> node --test) is out of scope here. Issue: #620 User-Visible: no